Graduated from the Royal (Dick) School of Veterinary Studies, Edinburgh University, in 1984, and gained a PhD in pancreatic and intestinal function at the University of Leicester in 1988. An internship at the University of Pennsylvania and residency in small animal medicine at THE Ohio State University were followed by a lectureship at Royal Veterinary College. He joined the faculty at Cornell University, Ithaca, NY in 1995. He is a past-president of the comparative gastroenterology society recipient of the National Phi Zeta and Pfizer awards for research, AVMF/AKC Career Achievement Award in Canine Research, the BSAVA Bourgelat Award for outstanding contributions to the field of small animal practice. His research interests are focused on inflammatory diseases of the GI tract (including the pancreas and liver), host bacterial interactions in health and disease, and culture independent bacteriology. Diplomate - American College of Veterinary Internal Medicine; Diplomate - European College of Veterinary Internal Medicine; Professor, Section of Small Animal Medicine; Professor of Veterinary Medicine in Medicine, Weill Cornell Medical College